Principal investigator

Amael Borzee_Hynobius sampling-square
Amaël Borzée – field work for Hynobius sp. in Korea

I am principally interested in behavioural ecology and the conservation of species. My focus is currently shifting away from exclusively studying amphibian breeding behaviour and conservation in North East Asia (summary of the projects here), although this will always remain one of my research strengths. I now wish to address broader questions aiming at the conservation of multi-species populations over large landscapes, including multiple types of approaches and analytical tools. Find my publications below.

My past projects mostly revolved around the reasons behind the endangered status of the Suweon Treefrog (Dryophytes suweonensis), the impact of climate change on the breeding phenology of Korean amphibians, and the eco-phylogeography of amphibians in North East Asia. Most results are available here.

My research interests have broadened since the establishment of my lab in September 2019, and you can find the lab page on Research Gate and Facebook here!

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Research is not (really) everything, and I very much enjoy looking up new species on my way around. I try identifying what I can, and most of it is on iNaturalist.

iNat obs
Map of my observations as of 05 August 2020
Amaël Borzée – New Zealand 2020 – for the 9th World Congress of Herpetology


Othman S. N, Chuang M-F., Kang H., Bae Y., Kim A., Jang Y. & Borzée A. (in press). Methodological guidelines for minimally invasive tail-clipping: a case study from Rana huanrenensis tadpoles. Asian Journal of Biological Conservation. in press

Shin Y. & Borzée A. (in press). Melanism in the Ussuri pitviper (Gloydius ussuriensis) from the Republic of Korea, with remarks on color variations in the species. Jordan Journal of Natural History. in press

Borzée A., McNeely J., Magellan K., Miller J. R. B., Porter L., Dutta T., Kadinjappalli K. P., … & Zhang L. (in press). COVID -19 triggered regulation of wildlife trade in Asia. Trends in Ecology & Evolution. DOI: 10.1016/j.tree.2020.10.001

Moores N. & Borzée A. (in press). First adequately-documented record of Chinese Leaf Warbler Phylloscopus yunnanensis in the Republic of Korea. Forktail.

Ambu J. & Borzée A. (in press). First record of putative necrophilia in Huanren brown frogs (Anura: Ranidae: Rana huanrenensis). Herpetology Notes. in press

Jowers M. J., Rivas G. A., Jadin F., Braswell A. L., Auguste R. J., Borzée A. & Murphy J. C. (in press). Assessing the monophyly of the widespread Black-headed Snake, Tantilla melanocephala in the Eastern Caribbean. Amphibian & Reptile Conservation.

Borzée A., Sin E., Oh S. & Jang Y. (2020) Spring voices in Korean rice fields: the effect of abiotic variables and syntopic calls on the calling activity of the treefrog Dryophytes suweonensis. Asian Herpetological Research. in press

Andersen A., Yi Y., Borzée A., Kim K., Moon K., Kim J., Kim T., & Jang Y. (in press). Modelling population dynamics and habitat connectivity for a reintroduced ursid population. The Oryx.

Son E. & Borzée A. (2020). Pelophylax chosenicus (Gold-spotted Pond Frog). Herpetological Review. 51(3):134

Messenger K., Dahn H., Andersen D., Shin Y., Yi Y., Bae Y. & Borzée A. (2020). Bufo stejnegeri (Korean Water Toad). Herpetological Review. 51(3):532

Son E., Bae Y., Park J. & Borzée A. (2020). Dryophytes suweonensis (Suweon Treefrog). Herpetological Review. 51(3): 532

Shin Y., Jang Y.,  Allain S. J. R. & Borzée A. (2020). Catalogue of herpetological specimens of the Ewha Womans University Natural History Museum (EWNHM), Republic of Korea. ZooKeys. 965: 103–139. DOI: 10.3897/zookeys.965.52976.

Borzée A. (2020). Recommendations for IUCN Red List Conservation Status of the “Dryophytes immaculatus Group” in North Eeast Asia. Diversity. 12:336. DOI: 10.3390/d12090336

Borzée A., Kwon S., Koo KS. & Jang Y. (2020). Policy recommendation on the restriction on amphibian trade towards the Republic of Korea.. Frontiers in Environmental Sciences. 8:129. DOI: 10.3389/fenvs.2020.00129

Othman S. N., Chen Y.-H., Chuang M.-F., Andersen D., Jang Y., Borzée A. Impact of the Mid-Pleistocene Revolution and anthropogenic factors on the dispersion of Asian black-spined toads (Duttaphrynus melanostictus). Animals. 10:1157. DOI: 10.3390/ani10071157

Borzée A., Andersen D., Groffen J., Kim Y. I., An J., Othman S., Ri K., Nam T. Y., Bae Y., Ren J-L., Li., J-T., Chuang M-F., Yi Y., Shin Y., Kwon T., Jang Y., & Min M-S. (2020). Yellow sea mediated segregation between North East Asian Dryophytes species. Plos One. 15(6): e0234299. DOI: 10.1371/journal.pone.0234299

Amin H., Borzée A & Messenger K. (2020). Rana zhenhaiensis (Zhenhai brown frog). Predation. Herpetological Review. 51(2): 304.

Litvinchuk S. N., Schepina N. A. & Borzée A. (2020). Reconstruction of past distribution for the Mongolian toad, Strauchbufo raddei (Anura: Bufonidae) using environmental modeling. PeerJ. 8:e9216. DOI: 10.7717/peerj.9216

Shin Y., Ambu J. & Borzée A. (2020). Additional observations on heterospecific amplexus in Asiatic toads (Anura: Bufonidae: Bufo gargarizans) in the Republic of Korea. Herpetology notes. 13:411-413.

Borzée A., Fong F. Nguyen Q. H. & Jang Y. 2020. Large-Scale Hybridisation as an Extinction Threat to the Suweon Treefrog (Hylidae: Dryophytes suweonensis). Animals 10(5):764. DOI: 10.3390/ani10050764

Borzée A., Nguyen H., & Jowers M. (2020). Advertisement and antagonistic call description of the endemic and enigmatic Trinidad Thin-toed Frog Leptodactylus nesiotus. South American Journal of Herpetology. 15:63-67. DOI: 10.2994/SAJH-D-18-00011.1

Kim M, Nguyen G. H., Yi Y., Ahn J., Kim E. U., Jang S., Kim S., Kong S., Borzée A. (2020) Policy recommendation on whaling, trade and watching of cetaceans in the Republic of Korea. Biodiversity Journal. 11 (1):255–258. DOI: 10.31396/Biodiv.Jour.2020.

Shin Y., Jeong D. & Borzée A. (2020). Mass displacement of the Korean clawed salamander (Onychodactylus koreanus) and the threat of road-kill. Herpetological bulletin.151:28-31. DOI: 10.33256/hb151.2831

Shin Y., Jang Y. & Borzée A. Gloydius ussuriensis (Ussuri Mamushi). (2020). Disease. Herpetological Review. 51(1): 144

Shin Y., Jang Y. & Borzée A. Gloydius saxatilis (Rock Mamushi). (2020). Dicephalism. Herpetological Review 51(1): 143-144.

Kim K., Macias D., Borzée A. and Jang Y. (2020). Ueno’s brown frog Rana uenoi indiscriminately ceases calling in the presence of daytime birds. Ethology Ecology and Evolution 32:251-263. DOI: 10.1080/03949370.2020.1717638

Marina M. T., Puan C. L., Chuang M-F., Othman S. N. & Borzée A. (2020). First record of ultraviolet fluorescence in Bent-toed Gecko Cyrtodactylus quadrivirgatus (Gekkonidae: Sauria). Herpetology notes 13:211-212.

Shin Y., Jang Y., Kim T. & Borzée A. (2020). A preserved specimen of Karsenia koreana (Caudata: Plethodontidae) misidentified and labelled as Hynobius leechii 27 years before the species’ description. Current Herpetology 39(1):75–79 DOI: 10.5358/hsj.39.75.

Shin Y., Jang Y. & Borzée A. (2020) Limb malformations in Bombina orientalis (Anura: Bombinatoridae) in the Republic of Korea based on museum specimens. Herpetology notes. 13: 29-31

Borzée A., Purevdorj Z., Kim Y. I., Kong S., Choe M., Yi Y., Kim K., Kim A. & Jang Y. (2019) Breeding preferences in the Treefrogs Dryophytes japonicus (Hylidae) in Mongolia. Journal of Natural History. 53:43-44 DOI: 10.1080/00222933.2019.1704458

Groffen J., Oh S-Y., Kwon S., Jang Y. & Borzée A. (2019). High mortality in Bufo gargarizans eggs associated with a non-described Saprolegnia ferax strand in the Republic of Korea. Diseases of Aquatic Organisms. 137:89-99. DOI: 10.3354/dao03434

Groffen J., Kong S., Jang Y. and Borzée A. (2019). The invasive American bullfrog (Lithobates catesbeianus) in the Republic of Korea: history and recommendation for population control. Management of Biological Invasions. 10(3):517-535. DOI: 10.3391/mbi.2019.10.3.08.

Borzée A., & Jang Y. (2019). Policy recommendation for the conservation of the Suweon Treefrog (Dryophytes suweonensis) in the Republic of Korea. Frontiers in Environmental Science. 7:39. DOI: 10.3389/fenvs.2019.00039.

Borzée A., Choi Y., Kim Y.E., Jablonski P. & Jang Y. (2019). Interspecific variation in seasonal migration and brumation behaviour in two closely related species of treefrogs. Frontiers in Ecology and Evolution. 7:55. DOI: 10.3389/fevo.2019.00055

Messenger R.K., Yi. Y. & Borzée A. (2019). UV biofluorescence in swallowtail butterfly larvae. Frontiers in Ecology and the Environment – EcoPics. 17(8):444. DOI: 10.1002/fee.2110

Bae Y., Kong S., Yi Y., Jang Y. & Borzée A. (2019). Additional threat to Hynobius salamander eggs: predation by loaches (Misgurnus sp.) in agricultural wetlands. Animal Biology. 69(4):451–461. DOI: 10.1163/15707563-20191070

Chuang M.-F., Choe M., Kang H., Borzée A., Kim A., Kwan S., Sung M., Jang Y. (2019). Microhabitat preference in American bullfrog tadpoles (Lithobates catesbeianus) in relation to predation pressure. Aquatic Invasions. (14):3 444–457. DOI: 10.3391/ai.2019.14.3.04

Borzée A., Andersen D., Groffen J., Kim H-Y., Bae Y. & Jang Y. (2019). Climate change-based models predict range shifts in the distribution of the only Asian plethodontid salamander: Karsenia koreana. Scientific reports. 9: 11838. DOI: 10.1038/s41598-019-48310-1

Borzée A., Ren J, Li J-T., Groffen J., Jang Y. & Messenger R.K. (2019). Microhabitat segregation between Black-spotted Pond Frogs, Pelophylax nigromaculatus, and Gold-striped Pond Frogs, P. plancyi (Anura: Ranidae). Reptiles & Amphibians: Conservation and Natural History. 26(2):119–120.

Borzée A., Baek H.J., Lee C.H., Kim D.Y., Song J-Y., Suh J-H., Jang Y. & Min M-S. (2019). Scientific publication of georeferenced molecular data as an adequate guide to delimit the range of Korean Hynobius salamanders through citizen science. Acta Herpetologica. 14(1):27-33. DOI: 10.13128/Acta_Herpetol-24102.

Chuang M-F. & Borzée A. (2019). Fejervarya limnocharis (Asian Rice Frog). Predation. Herpetological Review. 50(2):336.

Borzée A., Yi Y., Andersen D. K., Kim K., Moon K-S., Kim J.J., Kim T.W. & Jang Y. (2019). First dispersion events in reintroduced Asiatic Black Bear (Ursus thibetanus) in Korea. Russian Journal of Theriology. 18(1):51-55.

Chuang M-F., Borzée A. &, Kam Y-C. (2019). Female-female mating competition in Nidirana adenopleura (Ranidae, Nidirana) in Taiwan. Herpetology Notes. 12: 617-618.

Kim E., Nugraha Cahyana A., Jang Y. & Borzée A. (2019) Breeding range variation between Korean Hylids (Dryophytes sp.). Journal of Asia-Pacific Biodiversity. 12:135-138. DOI: 10.1016/j.japb.2018.12.002.

Heo K., Kim Y.I., Bae Y., Jang Y. & Borzée A. (2019). First report of Dryophytes japonicus tadpoles in saline environment. Russian Journal of herpetology. 26(2):87-90. DOI: 10.30906/1026-2296-2019-26-2-87-90

Murphy J.C, Santos J.L., Salvi D., Braswell A.L., Charles S.P., Borzée A. & Jowers M.J. (2019). The reduced limbed lizards of the genus Bachia (Reptilia, Squamata, Gymnophthalmidae); biogeography, cryptic diversity and morphological convergence in the eastern Caribbean. Organisms Diversity and Evolution. 19(2): 321–340. DOI: 10.1007/s13127-019-00393-4

Borzée A., Struecker M-Y., Yi Y., Kim H. & Kim D. (2019). Time for Korean wildlife conservation. Science. 363(6432):1161.2-1162. DOI: 10.1126/science.aaw9023

Borzée A. (2019). First amphibian behavioural observation from the Democratic People’s Republic of Korea: predation of a Dryophytes japonicus tadpole by Hydaticus sp. larvae. (2019). Nature Conservation Research. 4(1):106–108. DOI: 10.24189/ncr.2019.002

Murphy J.C., Braswel A.L., Charles S.P., Auguste R.J., Fuenmayor G.A.R., Borzée A., Lehtinen R.M. & Jowers M.J. (2019). A new species of Erythrolamprus from the oceanic island of Tobago (Squamata, Dipsadidae). ZooKeys. 817:131-157. DOI: 10.3897/zookeys.817.30811.

Groffen J., Borzée A. & Jang Y. (2019). First record of necrophilia in the prolonged breeder Rana uenoi. Herpetology Notes. 12:41-42.

Groffen J., Yang Y., Borzée A. & Jang Y. (2019). Interspecific amplexus between Glandirana tientaiensis (Chang, 1933) and Odorrana schmackeri (Boettger, 1892) at the Fuchun River, eastern China. Herpetology Notes. 12:41-42.

Chuang M-F., Borzée A. & Kam Y.C. (2018). Attendance to egg clutches by male Kurixalus eiffingeri increases hatching success and decreases predation through the recognition of invasive slugs (Parmarion martensi). Ethology. 125(1): 40-46. DOI: 10.1111/eth.12822.

Borzée A. Bae Y. & Seliger B. (2018). Pelophylax chosenicus (Gold-spotted Pond Frog). Herpetological Review. 49(4):707.

Borzée A. & Seliger B. (2018). Dryophytes suweonensis (Suweon Treefrog). Herpetological Review. 49(4):707.

Dufresnes C., Mazepa G., Rodrigues N., Brelsford A., Litvinchuk S. N., Sermier R., Betto-Colliard C., Blaser O., Borzée A., … & Jeffries D. L. (2018) Formal description of the new tree frog species inhabiting Northern Italy and Southern Switzerland. Bulletin de la Societe Vaudoise des Sciences Naturelles 97:5-11.

Maslova I., Jang Y., Zhestkov A. & Borzée A. (2018). Dryophytes japonicus (Japanese Treefrog). Colouration. Herpetology Notes. 11:1007-1008.

Borzée A., Yu A-Y., & Jang Y. (2018). Variations in boldness, behavioural and physiological traits of an endangered and a common hylid species from Korea. Ethology Ecology & Evolution. 30(6):515-533. DOI: 10.1080/03949370.2018.1441192

Jeffries L.D., Lavanchy G., Sermier R., Sredl M.J., Miura I., Borzée A., Barrow L.N., Canestrelli D., … & Perrin N. (2018). A rapid rate of sex-chromosome turnover and non-random transitions in true frogs. Nature Communications. 9(1):4088. DOI: 10.1038/s41467-018-06517-2.

Dufresnes C., Mazepa G., Rodrigues N., Brelsford A., Litvinchuk S. N., Sermier R., Betto-Colliard C., Blaser O., Borzée A., … & Jeffries D. L. (2018) Genomic evidence for cryptic speciation in tree frogs from the Apennine Peninsula, with description of Hyla perrini sp. nov. Frontiers in Ecology and Evolution. 6:144. 10.3389/fevo.2018.00144.

Borzée A., Kong S., Didinger C, Nguyen H.Q. & Jang Y. (2018). A ring-species or a ring of species? Phylogenetic relationship between two treefrog species around the Yellow Sea: Dryophytes suweonensis and D. immaculatus. (2018). The Herpetological Journal. 28:160-170.

Groffen J., Borzée A. & Jang Y. (2018). Rana uenoi (Prevernal frog). Predation. Herpetological Review. 49(3):519.

Bae Y., Borzée A., Heo J., & Jang Y. (2018). Rana uenoi (Ueno’s brown frog). Leech parasitism. Herpetological Review.49(3):519.

Borzée A., Andersen D. & Jang Y. (2018). Population trend inferred from aural surveys for calling anurans in Korea. PeerJ. 6:e5568; DOI 10.7717/peerj.5568.

Borzée A., Kim YI., Kim K. & Jang Y. (2018).  Methodological development for harmonic direction finder tracking in salamanders. Herpetological Conservation and Biology. 3(2):473-478.

Maslova I., Rodina E. E., Jang Y. & Borzée A.  (2018). Breeding populations of Bombina orientalis in degraded urban habitat in Vladivostok, Russia. Biodiversity Journal. 9(2): 127-130.

Macias D., Groffen J., Jang Y & Borzée A. (2018). Rana coreana (Korean brown frog) and R. uenoi (Ueno’s Brown Frog). Hibernacula. Herpetological Review. 49(2):307-308.

Groffen J., Borzée A. & Jang Y. (2018). Positioning of two treefrog species within rice paddies in relation to different habitat borders. Animal Cells and Systems. 22(3):205-211.

Borzée, A., Heo, K. & Jang, Y. (2018). Relationship between agro-environmental variables and breeding Hylids in rice paddies. Scientific reports. 8:8049. DOI: 10.1038/s41598-018-26222-w

Borzée A., Kim YI., Kim Ye & Jang Y. (2018). Translocation of the endangered and endemic Korean treefrog Dryophytes suweonensis. Conservation Evidence. 15:6-11.

Borzée A. & Jang Y. (2018). Interference competition driven by hydric stress in Korean Hylids. Nature Conservation Research. 3: suppl. 1:120-124.

Borzée A., Groffen J. & Seliger B. (2018). Bombina orientalis (Oriental Fire-bellied Toad). Herpetological Review. 49(1): 69.

Borzée A., Kim M., Kim JY., Kim T. & Jang Y. (2018). Microhabitat use during brumation in the Japanese treefrog, Dryophytes japonicus. 39:163-175. Amphibia-Reptilia. 39:163-175. DOI: 10.1163/15685381-17000036

Kim K., Kong S., Kim Y. I., Borzée A., Bae Y & Jang Y. (2018). Japanese Hard Ticks (Ixodes nipponensis) Parasitizing on the Endangered Leopard Cat (Prionaliurus bengalensis euptilurus) in the Republic of Korea. Animal Systematics, Evolution and Diversity. 34(1):23-26.

Borzée A., Kyong C.N., Kil HK. & Jang Y. (2018). Impact of water quality on the occurrence of two endangered Korean anurans: Dryophytes suweonensis and Pelophylax chosenicus. Herpetologica. 74(1):1-7.

Kim M.Y., Borzée A., Kim J.Y. & Jang Y. (2017). Lateral line as a mean of individual identification through visual and software assisted methodologies. Journal of Ecology and Environment. 41:42.

Borzée A., Santos J. L., Sanchez-Ramirez S., Bae Y., Heo K., Jang Y. & Jowers M. (2017). Phylogeographic and population insights of the Asian common toad (Bufo gargarizans) in Korea and China: population isolation and expansions as response to the ice ages. PeerJ. 5:e4044 DOI: 10.7717/peerj.4044

Borzée A. & Jang Y. (2017). Impact of rice and bean harvests on the Suweon Treefrog (Dryophytes suweonensis). International Journal of Current Research. 9(10):59620-59623.

Borzée A., Kim K., Heo K., Jablonski P. & Jang Y. (2017). Impact of reclamation and agricultural water regime on the distribution and conservation status of Dryophytes suweonensis. PeerJ. 5:e3872 DOI: 10.7717/peerj.3872

Nguyen H., Kim Y. I., Borzée A. and Jang Y (2017). Efficient isolation method for high-quality genomic DNA from cicada exuviae. Ecology and Evolution. 7:8161–8169. 10.1002/ece3.3398

Kim HT., Bae YH., Kim H., Kim D. & Borzée A. (2017). Gekko japonicus (Schlegel’s Japanese Gecko). Herpetological Review. 48(3):82.

Borzée A., Bae Y. H., Groffen J. & Jang Y. (2017). Pelophylax chosenicus (Gold-spotted Pond Frog). Herpetological Review. 48(3):79-80

Borzée A., Kosch T., Kim M.Y. & Jang Y. (2017). Introduced bullfrogs are associated with increased Batrachochytrium dendrobatidis prevalence and reduced occurrence of Korean treefrogs. Plos One. 12(5): e0177860. DOI: 10.1371/journal.pone.0177860

Borzée A., Yi Y., Kusrini M.D. & Jang Y. (2017). Habitat use by the Javan caecilian (Ichthyophis hypocyaneus). Korean Journal of Herpetology. 8: 15-18,

Borzée A, Didinger C, Jang Y. (2017). The Complete Mitochondrial Genome of Dryophytes suweonensis (Anura Hylidae). Mitochondrial DNA Part B: Resources. 1, 2: 5-6, DOI: 10.1080/23802359.2016.1275833.

Dufresnes C., Litvinchuk S., Borzée A., Jang Y., Li J-T., Miura I., Perrin N. & Stöck M. (2016). Phylogeography reveals an ancient cryptic radiation in East-Asian tree frogs (Hyla japonica group) and complex relationships between continental and island lineages. BMC Evolutionary Biology. 16:253, DOI: 10.1186/s12862-016-0814-x.

Borzée A., Kim JY., da Cunha M.A.M., Lee D., Sin E., Oh S., Yi Y. & Jang Y. (2016). Temporal and spatial differentiation in microhabitat use: Implications for reproductive isolation and ecological niche specification. Integrative Zoology. 11(5):375-387.

Borzée A., Kim JY. & Jang Y. (2016). Asymmetric competition over calling sites in two closely related treefrog species. Scientific Reports, 6:32569, DOI: 10.1038/srep32569

Borzée A., Kim K. & Jang Y. (2016). Rana uenoi (Prevernal Frog). Herpetological Review. 47, 3:421.

Borzée A., Yu SH. & Jang Y. (2016). Dryophytes suweonensis (Suweon Treefrog). Herpetological Review. 47, 3:418.

Borzée A., Park G., Kim S. & Jang Y. (2016). Korean long-tailed gorals (Naemorhedus caudatus) are rare but successfully reproducing in remote mountains. International Journal of Sciences: Basic and Applied Research, 27, 2:1-6

Borzée A., Kim JY., & Jang Y. (2016). Kaloula borealis (Boreal Digging Toad). Habitat use. Herpetological Review. 47(1).

Dufresnes C., Majtyka T., Baird D., Gerchen J., Borzée A., Savary R., Ogielska M., Perrin N. & Stöck M. (2016). Empirical evidence for large X-effects in animals with undifferentiated sex chromosomes. Scientific reports, 6:21029, DOI: 10.1038/srep21029.

Borzée A., Ahn J., Kim S., Heo K., & Jang Y. (2015). Seoul, keep your paddies! Implications for the conservation of hylid species. Animal Systematics, Evolution and Diversity, 31, 3:176-184.

Dufresnes C., Borzée A., Horn A., Stöck M., Ostini M., … & Perrin N. (2015) Sex-chromosome homomorphy in Palearctic tree frogs results from both turnovers and X-Y recombination. Molecular Biology & Evolution, msv113.

Borzée A., & Jang Y. (2015). Description of a semi-natural habitat of the endangered Suweon Treefrog, Hyla suweonensis. Animal Cells and Systems, 19, 3:1-5.

Jang Y. & Borzée A. (2014). Research on Microhabitat Differentiation between Two Treefrog Species May Reveal the Cause of Population Decline in the Endangered Hyla suweonensis in Korea. Froglog, 112:4, 48-50.

Roh G., Borzée A., & Jang Y. (2014). Spatiotemporal distributions and habitat characteristics of the endangered treefrog, Hyla suweonensis, in relation to sympatric H. japonica. Ecological Informatics, 24, 78-84.

Borzée A., Park S., Kim A., Kim HT., & Jang Y. (2013). Morphometrics of two sympatric species of tree frogs in Korea: a morphological key for the critically endangered Hyla suweonensis in relation to H. japonica. Animal Cells and Systems, 17(5), 348-356.

Stöck M., Dufresnes C., Litvinchuk SN., Lymberakis P., Biollay S., Berroneau Borzée A., … & Perrin N. (2012). Cryptic diversity among Western Palearctic tree frogs: Postglacial range expansion, range limits, and secondary contacts of three European tree frog lineages (Hyla arborea group). Molecular Phylogenetics and Evolution, 65(1), 1-9.